This research was supported by the Ajou University research fund (J-S.L.) and grants from the Institute for Basic Science ( IBS-R013-A1 ) and Basic Research Program through the Korea Brain Research Institute ( 20-BR-03-01 ) funded by the Korean Ministry of Science and ICT (S.C.) , and KAKENHI ( 17H04011 , 20H03409 , 18K06711 ) of the Ministry of Education, Culture, Sports, Science, and Technology (MEXT) , the Vehicle Racing Commemorative Foundation , Takeda Science Foundation , Koyanagi Foundation , Life Science Foundation of Japan , KOS\\u00C9 Cosmetology research Foundation , and The Uehara Memorial Foundation (T.F., T.H.) . This work was also supported by the National Research Foundation of Korea (NRF) grant funded by the Korean government (MSIT) ( 2019005607 ) (B-H.B.).